Bulbul, Greenbul, Brownbul, Leaflove, Bristlebill
Order Passeriformes is the largest order of birds in the Class Aves. It contains over 140 families with 6.500 identified bird species world wide. They are also called perching birds. 3 toes point forward, 1 backwards called hallux and is long. This is called anisodactyl arrangement. This toe constellation makes it easier to hold on to a branch, twig, wire or blade of grass.
A further adaptation for perching is a tendon running on the rear side of the leg to the underside of the toes is attached to the muscle behind the Tibiotarsus (the large bone between femur and tarsometatarsus). This makes the feet curl automatically once the leg is bent and becomes stiff when landing on a branch. So they can sleep without falling down and using valuable energy. Bird foot
Passerines are mostly insectivorous or omnivorous meaning feeding on insects, small vertebrates, fruits, seeds and nectar. Omnivorous birds change to a more carnivorous diet during breeding season. Plant material has less protein which is required for the chicks to grow fast and strong.
The name Passeriformes derives from Greek – Passer = Sparrow, Formis = shape. They originated in the Southern Hemisphere around 60 Mio years ago.
Most Passerines have 12 tail feathers. The eggs are coloured in most species. The number of eggs vary from species to species. The chicks are altricial meaning they hedge blind, without feathers and are helpless. They have reflexes to sense when a parent is landing and open their beak wide to get fed. The beak is fringed with yellow skin to signal the feeding parent where to place food.
In the Family Pycnonotidae are Bulbuls, Greenbuls, Brownbuls, Leafloves and Bristlebills
166 species in 32 genera are spread from Africa as far as Japan.
Bulbuls and other members of the same family are slender birds with short necks, long tails and short rounded wings. The colour varies from grey to brown, greenish, yellow. The eye colour is from dark to yellow, while the rings around the eyes are also variable from species to species. Some species can raise their neck feathers to impress others.
Majority of Bulbuls are omnivorous. They feed on insects, fruits, seeds, nectar and even small vertebrates. Most feed dominantly on fruits and seeds and supplement food with insects. I often observe the Dark-capped Bulbuls in my garden hunting butterflies in flight or catching different insects on the ground.
Members of the family Pycnonotidae are monogamous. The exeption is the Yellow-whiskered Greenbul. It appears to be polygamous and engages in lekking. This means several males gather at one place and display in competition. The females observe these males and chose their partner.
Most Bulbul species are quite tolerant to the habitat. They can thrive in disturbed areas, gardens, open areas and forest edges. Some species living in rain forests are under pressure from pet trade and habitat destruction.
Greenbuls like leaved forest and are not always easy to spot.
Many species are split into subspecies. Depending on the author some are treated as full species. Changes are made every now and then.
To give you as actual information as possible we use Avibase, the books “Birds of Africa South of the Sahara”, “Birds of East Africa”, “Birds of Kenya and Northern Tanzania”. Then we put the most characteristic information to the photo.
You can also follow us on Facebook “Bird photography Safaris Kenya” and see the numerous species as well as the beautiful photos from Lorenzo Barelli. Click on the photo for larger view. The names of the photographers are on the photo and in the text. Without the generosity of several amazing bird photographers it had not been possible to create the pages with so many illustrated bird species. Those photos without name are from Elvira Wolfer.
Here you find samples of our birding safaris. Some below listed species are endemic and one needs to know the exact locations where to find them. We know the spots and can generate the journey to see several of these special Greenbuls, Brownbuls and Leaflove species.
Dark-capped Bulbul, Graubülbül (Pycnonotus tricolor) is common in various biotopes even gardens from E Cameroon, South Sudan, E to Southern Africa.
Yellow-streaked Greenbul or Yellow-streaked Bulbul (Phyllastrephus flavistriatus tenuirostris) lives in forest from NE Tanzania to NE Mozambique. There has been no sighting in SE Kenya since 1938. Photo Per Holmen, Amani Forest
Yellow-streaked Greenbul or Yellow-streaked Bulbul (Phyllastrephus flavistriatus olivaceogriseus) is found in highland forest from E DRC to SW Uganda. Photo Per Holmen, Bwindi impenetrale Forest
Other subspecies P. f. kungwensis from Kungwe-Mahari Mountains, W Tanzania, P. f. uzungwenis in Uzungwe Mountain, E Tanzania.
White-throated Greenbul or White-throated Bulbul (Phyllastrephus albigularis) is found in various types of forests in Uganda and farther West. Photo Per Holmen, Bundungo Forest
Icterine Greenbul (Phyllastrephus icterinus) is more a W African species but can be found in far W of Uganda from 700 – 1.250 m in undergrowth and primary forest.
Xavier’s Greenbul (Phyllastrephus x. xavieri) lives in far W Uganda and prefers tangled vines and creepers in midstoreys.
Cabanis’s Greenbul or Cabanis Bulbul (Phyllastrephus cabanisi sucosus) is the subspecies in W Kenya, Rwanda, Uganda, NW Tanzania. Small noisy flocks move in dense undergrowth. Photo Per Holmen, Kakamega Forest
Other subspecies: P. c. nandensis from the Northern Nandi Hills, P. c. ngurumanensis from the Nguruman mountain – (both not recognised by all authors).
Placid Greenbul (Phyllastrephus placidus) is a split from Cabanis’s Bulbul. It prefers undergrowth in mid to high altitude forest. Photo Per Holmen
Fischer’s Greenbul (Phyllastrephus fischeri) is a coastal bird – 600 m in forest undergrowth from SE Somalia, Kenya, Tanzania to N Mozambique. The are more often heard than seen. Photo Per Holmen, East Usambara
Grey-olive Greenbul or Grey-olive Bulbul (Phyllastrephus cerviniventris) keeps to dense, tangled vegetation on or near water below 1.700 m from S Kenya to Tanzania and farther South. There is a population around Mt.Kenya. Photo Per Holmen
Lowland tiny Greenbul (Phyllastrephus debilis rabei) they are uncommon in undergrowth in pairs or small flocks in coastal forest from Kenya to Rufiji River in Tanzania. Photo Per Holmen, Arabuko Sokoke Forest. The nominate form occurs from SE Tanzania to Mozambique.
Toro olive Greenbul (Phyllastrephus hypchloris) they live in pairs or small parties in entangled vegetation in prime forest from 700 – 1.800 m in DRC, Uganda, W Kenya and far NW Tanzania. They are shy and uncommon.
Usambara Greenbul or Montane tiny Greenbul (Phyllastrephus albigula) is endemic to the Nguru and Usambara Mountains from 300 – 1.500 m. Near threatened due to habitat / forest destruction.
Sassi’s Greenbul (Phyllastrephus lorenzi) occurs in a small range in primary forest of E DRC and adjacent Uganda.
Northern Brownbul or East African Scrub Bulbul (Phyllastrephus strepitans) is endemic to NE Africa from 0 – 1.000 m. It is a species of drier areas with less tangled vegetation. Tsavo East National Park
Terrestrial Brownbul (Phyllastrephus terrestris suahelicus) small flocks are found in coastal forests of Kenya and Tanzania to N Mozambique. 2 populations are in low altitude inland. Photo Lorenzo Barelli
Little Greenbul (Eurillas v. virens) is patchly distributed in thick secondary growth mainly below 1.500 m in Burundi, Uganda, SW Kenya. Photo Per Holmen, Kibale Forest
Little Greenbul (Eurillas virens zombensis) occurs in DRC, E Kenya, from NE to SW Tanzania. They are shy and often more heard than seen. Photo Per Holmen, Amani Forest. Zanzibar island has its own subspecies – E. v. zanzibarica.
Plain Greenbul (Eurillas c. curvirostris) lives in primary and secondary forest from 700 – 2.300 m from W Africa to Uganda and W Kenya. They are locally common in undergrowth. Photo Per Holmen, Kakamega Forest
Little grey Greenbul (Eurillas gracilis ugandae) prefers mid canopy and forest edge in Uganda where it locally common, uncommon in Kakamega Forest and far NW Tanzania. Photo Per Holmen, Minziro Forest
Yellow-whiskered Greenbul (Eurillas l. latirostris) inhabits forests and gardens from 700 – 3.000 m from Central Kenya to Uganda. Photo Elvira Wolfer, Aberdare at The Ark. The subspecies E. l. australis is found on the Ufipa Plateau, SW Tanzania
Ansorge’s Greenbul (Eurillas ansorgei kavirondensis) is known from East Africa in Kakamega, Kavirondo North and Mount Elgon. The subspecies E. a. ansorgei is found in Uganda only in Bwindi impenetrable Forest and then more in Western Africa. Photo Per Holmen, Kakamega Forest
Ansorge’s Greenbul (Eurillas ansorgei kavirondensis) is known from East Africa in Kakamega, Kavirondo North and Mount Elgon. The subspecies E. a. ansorgei is found in Uganda only in Bwindi impenetrable Forest and then more in Western Africa. Photo Per Holmen, Kakamega Forest
Sombre Greenbul (Andropadus importunus insularis) is more common along the EA coast than inland in forests and thickets. Photo Per Holmen, Tanga. The subspecies A. i. hypoxanthos lives in SE Tanzania.
Slender-billed Greenbul (Stelgidillas g. gracillirostris) is a common forest species from 700 – 2.500 m in Uganda and Western Kenya. Photo Per Holmen, Kakamega Forest
Slender-billed Greenbul (Stelgidillas gracillirostris percivali) is restricted to Kenya highland forests. Photo Per Holmen, Castle Forest Lodge, Mt. Kenya
Shelley’s Greenbul (Arizelocichla masukuensis roehli) lives in singles and pairs in mainly interior highland forest from 500 – 2.300 m from NE to S Tanzania. Photo Per Holmen, Amani Forest. The nominate A. m. masukuensis is endemic to Mt. Rungwe, SW Tanzania
Kakamega Greenbul (Arizelocichla k. kakamegae) is endemic to E DRC, Uganda (Albertine Rift) and W Kenya. Its special behaviour is moving slowly along mossy tree trunks and branches searching for insects. Photo Per Holmen, Kakamega Forest. The subspecies A. k. kungwensis is endemic to Kungwe-Mahari Mountains W Tanzania.
Kikuyu Mountain Greenbul or Olive-breasted Mountain Greenbul (Arizelocichla kikuyuensis) is a highland forest species from E DRC, Uganda, Rwanda and Central Kenya. Photo Lorenzo Barelli, Aberdare
Eastern Mountain Greenbul (Arizelocichla nigriceps nigriceps) is in highland forests of S Kenya (Nguruman Mountains) to N Tanzania. Photo Per Holmen, Karatu.
Arizelocichla nigriceps kungwensis occurs at Kungwe Mountain, Kigoma. Arizelocichla nigriceps usambarae is found from Taita Hills to Usambara Mountains.
Black-browed Mountain Greenbul (Arizelocichla fusciceps) inhabits mountains in SW Tanzania and adjacent countries.
Uluguru Mountain Greenbul (Arizelocichla neumanni) is endemic in the Uluguru Mountains in Tanzania. Photo Per Holmen, Uluguru Mountains
Yellow-throated (Mountain) Greenbul (Arizelocichla chlorigula) is endemic in E Tanania, Nguru Mountains and highlands of Iringa Distrikt. Photo Per Holmen, Mufindi
Yellow-bellied Greenbul (Chlorocichla flaviventris centralis) is found in forests with thick covers from 0 – 2.100 m from Somalia, Kenya to E + N Tanzania. Photo Elvira Wolfer, Lake Manyara
C. f. occidentalis occurs in W Tansania, DRC, Angola to Southern Africa.
Joyful Greenbul or Joyful Bulbul (Chlorocichla l. laetissima) is a very vocal bird in forests with high rainfall in W Kenya and W Uganda to DRC. Photo Lorenzo Barelli, Kakamega Rainforest